Our Leadership
At Greenstone Biosciences, our team is a convergence of world-class scientists, visionary technologists, and experienced drug development leaders. Co-founded by Dr. Joseph C. Wu, a trailblazer in iPSC and organoid research, our multidisciplinary group brings together decades of expertise from institutions like Stanford, Harvard, and UCLA.
With deep roots in New Approach Methods (NAMs), computational biology, and translational science, we are redefining drug discovery—advancing safer, more effective therapies with speed, precision, and human relevance.
United by purpose, driven by data, and focused on impact, the Greenstone team is committed to transforming the future of medicine.
Our Team
Leadership
Joseph Wu, MD, PhD
CO-FOUNDER
Joseph Wu, MD, PhD
CO-FOUNDER
Jade Chao, MPH, JD
CO-FOUNDER
Jade Chao, MPH, JD
CO-FOUNDER
Lip-Bu Tan
Chair, Board of Directors & CEO, Intel Corporation
Lip-Bu Tan
Chair, Board of Directors & CEO, Intel Corporation
Amarjit Gill
Board Member
Amarjit Gill
Board Member
Scientific Advisory Board
Axel Brunger, PhD
SCIENTIFIC ADVISORY BOARD
Axel Brunger, PhD
SCIENTIFIC ADVISORY BOARD
James Zou, PhD
SCIENTIFIC ADVISORY BOARD
James Zou, PhD
SCIENTIFIC ADVISORY BOARD
Scientists
Mukhtar Ahmed, PhD
DIRECTOR, PROJECT & PORTFOLIO STRATEGY
Mukhtar Ahmed, PhD
DIRECTOR, PROJECT & PORTFOLIO STRATEGY
At Vanderbilt University, he advanced to Research Assistant Professor. He led studies on polarized membrane trafficking and vesicle tethering, applying quantitative cell biology, targeted proteomics, and live-cell single-molecule imaging to reveal how protein complexes assemble and regulate epithelial organization. His work has been published in leading journals and contributed to advancing methodologies in the field.
Mukhtar’s experience extends beyond the bench. He has guided cross-functional programs integrating discovery biology, preclinical development, and strategy, ensuring that innovative science is translated into clinically meaningful outcomes. He has also completed executive education at Wharton, equipping him with the business acumen to align scientific, regulatory, and operational priorities in a rapidly evolving biotech landscape.
Paul Pang, PhD
DIRECTOR, RESEARCH & DEVELOPMENT
Paul Pang, PhD
DIRECTOR, RESEARCH & DEVELOPMENT
Paul received his bachelor’s degree from the University of Maryland and completed his graduate thesis on kidney and vascular stem cells at Harvard Medical School. He then spent a year working at the University of Southern California to continue his research on stem cell biology and regenerative medicine before subsequently receiving his Ph.D. in Integrative Molecular & Biomedical Sciences from Baylor College of Medicine, where he was the recipient of the Claude W. Smith Fellowship Award for Outstanding Research Performance and the Dean’s Award of Excellence. Paul then completed his postdoctoral training at Stanford University in the lab of Dr. Joseph Wu, where he continued pursuing his curiosity and longstanding interest in precision health and drug discovery using human pluripotent stem cells for translational medicine.
For his work in disease modeling and drug discovery, Paul received numerous competitive awards, including the NIH Ruth L. Kirschstein F31 and F32 National Research Service Awards, the R21 Exploratory/Developmental Research Award, and the R44 Small Business Innovation Research Award. In addition to the NIH, Paul is also a recipient of the Idea Development Award from the DOD, the Discovery Quest Award and the Translational Research Award from the California Institute for Regenerative Medicine (CIRM), and other national and international awards from the American Heart Association (AHA) and the Muscular Dystrophy Association (MDA).
Todd Herron, PhD
DIRECTOR, hiPSC BIOBANK
Todd Herron, PhD
DIRECTOR, hiPSC BIOBANK
Souhrid Mukherjee, PhD
SENIOR SCIENTIST - AI & COMPUTATIONAL BIOLOGY
Souhrid Mukherjee, PhD
SENIOR SCIENTIST - AI & COMPUTATIONAL BIOLOGY
Jeremy Leitz, PhD
SENIOR SCIENTIST
Jeremy Leitz, PhD
SENIOR SCIENTIST
Evgenios Neofytou, MD
SENIOR SCIENTIST
Evgenios Neofytou, MD
SENIOR SCIENTIST
Ibrahim Ibrahim, PhD
SENIOR SCIENTIST
Ibrahim Ibrahim, PhD
SENIOR SCIENTIST
David Dolivo, PhD
PROJECT SCIENTIST
David Dolivo, PhD
PROJECT SCIENTIST
Angelos Oikonomopoulos, PhD
SENIOR SCIENTIST
Angelos Oikonomopoulos, PhD
SENIOR SCIENTIST
At Greenstone Biosciences, he champions the application of scalable hiPSC-derived cardiomyocyte platforms to accelerate precision drug discovery for cardiovascular disease. In addition to leading internal R&D initiatives, he manages cross-functional collaborations with academic and industry partners—aligning scientific goals, timelines, and deliverables to drive shared success. His work bridges cutting-edge stem cell technologies with therapeutic innovation, enabling partners to de-risk development and unlock new indications through human-relevant platforms.
Researchers
Chris Yan
ASSOCIATE DIRECTOR, COMMERCIAL STRATEGY & OPERATIONS
Chris Yan
ASSOCIATE DIRECTOR, COMMERCIAL STRATEGY & OPERATIONS
Celine Lai
ASSOCIATE SCIENTIST
Celine Lai
ASSOCIATE SCIENTIST
Sushma Shenoy, MS
ASSOCIATE SCIENTIST
Sushma Shenoy, MS
ASSOCIATE SCIENTIST
Rebecca Yu
SENIOR RESEARCH ASSOCIATE
Rebecca Yu
SENIOR RESEARCH ASSOCIATE
Yang Zhou, MS
RESEARCH ASSOCIATE
Yang Zhou, MS
RESEARCH ASSOCIATE
Parker Walther
RESEARCH ASSOCIATE
Parker Walther
RESEARCH ASSOCIATE
Zachary Cook
SENIOR RESEARCH ASSOCIATE
Zachary Cook
SENIOR RESEARCH ASSOCIATE
Consultants
Audrey Jia, MD, PhD
CONSULTANT
Audrey Jia, MD, PhD
CONSULTANT
ex-FDA Drug Quality Reviewer
Keith Zhang, MD, PHD
CONSULTANT
Keith Zhang, MD, PHD
CONSULTANT
Daniel Levy, PhD
CONSULTANT
Daniel Levy, PhD
CONSULTANT
DEL BioPharma LLC
www.delbiopharma.com
Yong Wang, PhD
CONSULTANT
Yong Wang, PhD
CONSULTANT
With 11 years of service at the U.S. FDA's Office of New Drug Products (ONDP), Dr. Wang led comprehensive CMC reviews for INDs, NDAs, ANDAs, and DMFs while participating in pre-approval inspections and providing strategic regulatory guidance during critical milestone meetings. His FDA tenure included extensive collaboration with clinical divisions to identify and resolve approvability issues through the application of regulatory science, ICH guidelines, and risk-based assessments.
Dr. Wang's industry expertise spans an additional 11 years at leading pharmaceutical companies including Roche, Novartis, and Johnson & Johnson, where he specialized in late-stage development and global regulatory strategy. Following his FDA service, he provided regulatory and CMC consulting for over five years to prominent organizations including Gilead, Cambrex, and Jiangsu Hengrui. His unique combination of regulatory, industry, and consulting experience positions Dr. Wang as a valuable advisor in navigating complex pharmaceutical development and regulatory pathways.
John Higgins, PhD
CONSULTANT
John Higgins, PhD
CONSULTANT
HR and Finances
Amit Parikh, MBA
CFO / OUTFORCE, LLP
Amit Parikh, MBA
CFO / OUTFORCE, LLP
Sanjay Vrajlal
SENIOR FINANCE MANAGER
Sanjay Vrajlal
SENIOR FINANCE MANAGER
Audra Soltis
HR MANAGER / OUTFORCE, LLP
Lingyun Xu, PhD
LAB MANAGER
Lingyun Xu, PhD
LAB MANAGER